G. M. Smith, K. Ruebens, S. Gaudzinski-windheuser, and T. E. Steele, Subsistence strategies throughout the African Middle Pleistocene: Faunal evidence for behavioral change and continuity across the Earlier to Middle Stone Age transition, J. Hum. Evol, vol.127, pp.1-20, 2019.

L. Bahram, P. F. Mitchell, and . Africans, African Archaeology from the Earliest Tool Makers to Most Recent Foragers, 2008.

M. Chazan, Radiometric dating of the Earlier Stone Age sequence in Excavation I at Wonderwerk Cave, South Africa: preliminary results, J. Hum. Evol, vol.55, pp.1-11, 2008.

F. Berna, Microstratigraphic evidence of in situ fire in the Acheulean strata of Wonderwerk Cave, Northern Cape province, South Africa, Proc. Natl. Acad. Sci, vol.109, pp.1215-1220, 2012.

J. Brink, S. Holt, and L. K. Horwitz, The Oldowan and Early Acheulean Mammalian Fauna of Wonderwerk Cave (Northern Cape Province, South Africa). African Archaeol. Rev, vol.33, pp.223-250, 2016.

K. Kuman, The earlier Stone Age in South Africa: site context and the influence of cave studies. In Breathing life into fossils: Taphonomic studies in Honor of, pp.181-198, 2007.

F. L. Forrest, Journal of Archaeological Science: Reports Zooarchaeological reconstruction of newly excavated Middle Pleistocene deposits from Elandsfontein, South Africa, J. Archaeol. Sci. Reports, vol.17, pp.19-29, 2018.

B. Asfaw, The earliest Acheulean from Konso-Gardula, Nature, vol.360, pp.732-735, 1992.

C. J. Lepre, An earlier origin for the Acheulian, Nature, vol.477, pp.82-85, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00648859

G. Suwa, H. Nakaya, and B. Asfaw, The Konso Formation Paleontological Assemblages: Collecting and Documentation Methodologies, pp.5-9, 2014.

A. Echassoux, Comportements de subsistance et modifications osseuses à l'aube de l' Acheuléen à Konso, Éthiopie. Anthropologie, vol.116, pp.291-320, 2012.

F. Diez-martín, The Origin of the Acheulean: The 1.7 Million-Year-Old Site of FLK West, Olduvai Gorge (Tanzania). Sci. Rep, vol.5, pp.1-9, 2015.

S. Semaw, The Early Acheulean ~1.6-1.2 Ma from Gona, Ethiopia, The Emergence of the Acheulean in East Africa and Beyond, pp.115-128, 2018.

R. Gallotti, An older origin for the Acheulean at Melka Kunture (Upper Awash, Ethiopia): Techno-economic behaviours at Garba IVD, J. Hum. Evol, vol.65, pp.594-620, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01842796

I. Fiore and A. Tagliacozzo, Taphonomic analysis of the bone remains from the Oldowan site of Garba IV, Studies on the Early Paleolithic Site of Melka Kunture, pp.639-682, 2004.

M. Domínguez-rodrigo, J. Serrallonga, L. Luque, F. Diez-martín, P. Bushozi et al., The archaeology of the Acheulean sites from South Escarpment, Peninj. A Research Project on the Archaeology of Human Origins, pp.205-226, 1995.

M. Domínguez-rodrigo, J. Serrallonga, L. Luque, F. Diez-martín, P. Bushozi et al., The archaeology of the North Escarpment, Peninj. A Research Project on the Archaeology of Human Origins, pp.227-256, 1995.

F. Diez-martín, Early Acheulean technology at Es2-Lepolosi (ancient MHS-Bayasi) in Peninj (Lake Natron, Tanzania), Quat. Int, pp.209-236, 2014.

M. Domínguez-rodrigo, Unraveling hominin behavior at another anthropogenic site from Olduvai Gorge (Tanzania): new archaeological and taphonomic research at BK, Upper Bed II, J. Hum. Evol, vol.57, pp.260-283, 2009.

M. Domínguez-rodrigo, On meat eating and human evolution: A taphonomic analysis of BK4b (Upper Bed II, Olduvai Gorge, Tanzania), and its bearing on hominin megafaunal consumption, Quat. Int, pp.129-152, 2014.

D. R. Braun, Early hominin diet included diverse terrestrial and aquatic animals 1.95 Ma in East Turkana, Kenya. Proc. Natl. Acad. Sci, vol.107, pp.10002-10007, 2010.

B. L. Pobiner, M. J. Rogers, C. M. Monahan, and J. W. Harris, New evidence for hominin carcass processing strategies at 1.5 Ma, Koobi Fora, Kenya, J. Hum. Evol, vol.55, pp.103-130, 2008.

S. R. Merritt, Investigating hominin carnivory in the Okote Member of Koobi Fora, Kenya with an actualistic model of carcass consumption and traces of butchery on the elbow, J. Hum. Evol, vol.112, pp.105-133, 2017.

R. L. Hay, Geology of the Olduvai Gorge, 1976.

G. L. Isaac and B. Isaac, Koobi Fora Research Project, Plio-Pleistocene Archaeology, 1997.

H. Roche, Les sites archéologiques plio-pléistocènes de la formation de Nachukui, Comptes Rendus -Palevol, vol.2, pp.663-673, 1997.

J. Quade, The Geology of Gona, Afar, Ethiopia. Geol. Soc. Am. Bull, vol.446, pp.1-31, 2008.

I. De-la-torre, The Early Stone Age lithic assemblages of Gadeb (Ethiopia) and the developed Oldowan/early Acheulean in East Africa, J. Hum. Evol, vol.60, pp.768-812, 2011.

Y. Beyene, The characteristics and chronology of the earliest Acheulean at Konso, Ethiopia. Proc. Natl. Acad. Sci, vol.110, pp.1584-1591, 2013.

R. Gallotti and M. Mussi, The Emergence of the Acheulean in East Africa: Historical Perspectives and Current Issues, The Emergence of the Acheulean in East Africa and Beyond (eds. Gallotti, R. & Mussi, M, pp.1-12, 2018.

A. Delagnes, Interpreting pachyderm single carcass sites in the African Lower and Early Middle Pleistocene record: A multidisciplinary approach to the site of Nadung'a 4 (Kenya), J. Anthropol. Archaeol, vol.25, pp.448-465, 2006.
URL : https://hal.archives-ouvertes.fr/halshs-00110608

F. Altamura, Archaeology and ichnology at Gombore II-2, Melka Kunture, Ethiopia: everyday life of a mixed-age hominin group 700,000 years ago, Sci. Rep, vol.8, pp.1-11, 2018.

F. Altamura, S. Gaudzinski-windheuser, R. T. Melis, and M. Mussi, Reassessing Hominin Skills at an Early Middle Pleistocene Hippo Butchery Site: Gombore II-2 (Melka Kunture, Upper Awash valley, Ethiopia), J. Paleolit. Archaeol, 2019.

M. C. Pante, The larger mammal fossil assemblage from JK2, Bed III, Olduvai Gorge, Tanzania: Implications for the feeding behavior of Homo erectus, J. Hum. Evol, vol.64, pp.68-82, 2013.

L. Balout, Préhistoire de l'Afrique du Nord. Essai de chronologie. (Arts et métiers graphiques, 1955.

R. Vaufrey, Préhistoire de l'Afrique, 1955.

P. Biberson, Le Paléolithique inférieur du Maroc atlantique, 1961.

J. D. Clark, The earlier Stone Age/Lower Palaeolithic in North Africa and the Sahara, New light on the northern African past, pp.17-37, 1992.

D. J. Mattingly, T. Reynolds, and J. N. Dore, Synthesis of human activities in the Fazzan, The Archaeology of Fazzan, pp.327-375, 2003.

T. Reynolds, The importance of Saharan lithic assemblages, Environment, Climate and Resources of the Libyan Sahara, pp.81-90, 2006.

L. Boudad and S. Guislain, Acquisition de supports prédéterminés destinés à la réalisation de bifaces: l' exemple de sites de surfaces du Sud-Est marocain, Anthropologie, vol.116, pp.364-377, 2012.

F. Parenti, D. Mengoli, and L. Natali, The Stone Age in Northwestern Libya: Observations Along a Pipeline, African Archaeol. Rev, vol.32, pp.413-441, 2015.

C. Denys, M. Patou, and N. Djemmali, Premiers résultats concernant l' origine de l'accumulation du matériel osseux de ce gisement Pléistocène, Comptes Rendus l'Academie Sci. -Ser. II, vol.299, pp.481-486, 1984.

D. Geraads, The Pleistocene Hominid site of Ternifine, Algeria: New results on the environment, age and human industries, Quat. Res, vol.25, pp.380-386, 1986.

J. Raynal, F. Sbihi-alaoui, D. Geraads, L. Magoga, and A. Mohib, The earliest occupation of North-Africa: the Moroccan perspective, Quat. Int, vol.75, pp.65-75, 2001.
URL : https://hal.archives-ouvertes.fr/halshs-00005359

J. P. Raynal, Hominid Cave at Thomas Quarry I (Casablanca, Morocco): Recent findings and their context, Quat. Int, pp.369-382, 2010.
URL : https://hal.archives-ouvertes.fr/halshs-00475759

C. Daujeard, D. Geraads, R. Gallotti, A. Mohib, and J. Raynal, Carcass Acquisition and Consumption by Carnivores and Hominins in Middle Pleistocene Sites of Casablanca (Morocco), J. Taphon, vol.10, pp.3-4, 2012.
URL : https://hal.archives-ouvertes.fr/hal-02377897

C. Daujeard, Pleistocene hominins as a resource for carnivores: A c. 500,000-year-old human femur bearing tooth-marks in North Africa, PLoS One, vol.11, pp.1-22, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01323864

D. Richter, The age of the hominin fossils from Jebel Irhoud, Morocco, and the origins of the Middle Stone Age, Nature, vol.546, pp.293-296, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01980421

J. Raynal, La grotte des Rhinocéros (Carrière Oulad Hamida 1, anciennement Thomas III, Casablanca), nouveau site acheuléen du Maroc atlantique, Comptes Rendus l'Académie des Sci. II, vol.316, pp.1477-1483, 1993.

E. Rhodes, J. Raynal, D. Geraads, and F. Sbihi-alaoui, Premières dates RPE pour l' Acheuléen du Maroc atlantique (Grotte des Rhinocéros, Casablanca), Comptes Rendus l'Academie Sci. -Ser. II, vol.319, pp.1109-1115, 1994.

E. J. Rhodes, J. S. Singarayer, J. P. Raynal, K. E. Westaway, and F. Z. Sbihi-alaoui, New age estimates for the Palaeolithic assemblages and Pleistocene succession of Casablanca, Morocco. Quat. Sci. Rev, vol.25, pp.2569-2585, 2006.
URL : https://hal.archives-ouvertes.fr/halshs-00092016

J. Raynal and A. Mohib, Préhistoire de Casablanca. I -La Grotte des Rhinocéros (fouilles 1991 et 1996), 2016.

D. Lefèvre and J. P. Raynal, Les formations plio-pléistocènes de Casablanca et la chronostratigraphie du Quaternaire marin du Maroc revisitées / The Plio-Pleistocene formations of Casablanca and the marine Quaternary chronostratigraphy of Morocco revisited, Quaternaire, vol.13, pp.9-21, 2009.

D. P. Geraads and . Carnivora, Mammalia) from Tighennif (Ternifine), vol.49, pp.445-458, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01393484

M. Domínguez-rodrigo, R. Barba, C. P. Egeland, and . Deconstructing-olduvai, , 2007.

M. Domínguez-rodrigo, Hunting and scavenging by the early hominids: the state of the debate, J. World Prehistory, vol.16, pp.1-53, 2002.

M. Domínguez-rodrigo and T. R. Pickering, Early Hominid Hunting and Scavenging: A Zooarcheological Review, Evol. Anthropol, vol.12, pp.275-282, 2003.

H. T. Bunn, Systematic Butchery by Plio/Pleistocene Hominids at Olduvai Gorge, Tanzania

, Curr. Anthropol, vol.27, pp.431-452, 1986.

M. Sahnouni, The first evidence of cut marks and usewear traces from the Plio-Pleistocene locality of El-Kherba (Ain Hanech), Algeria: Implications for early hominin subsistence activities circa 1.8 Ma, J. Hum. Evol, vol.64, pp.137-150, 2013.

M. Sahnouni, 1.9-million-and 2.4-million-year-old artifacts and stone tool-cutmarked bones from Ain Boucherit, Algeria. Science, vol.362, pp.1297-1301, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02173435

R. Potts and P. Shipman, Cutmarks made by stone tools on bones from Olduvai Gorge, Tanzania, Nature, vol.291, 1981.

H. T. Bunn, Early Pleistocene hominid foraging strategies along the ancestral Omo River at Koobi Fora, Kenya, J. Hum. Evol, vol.27, pp.247-266, 1994.

S. De-juana, A. B. Galán, and M. Domínguez-rodrigo, Taphonomic identification of cut marks made with lithic handaxes: An experimental study, J. Archaeol. Sci, vol.37, pp.1841-1850, 2010.

S. Schlanger, Recognizing persistent places in Anasazi Settlement systems, pp.91-112, 1992.

C. A. Tryon, Sites on the landscape: Paleoenvironmental context of late Pleistocene archaeological sites from the Lake Victoria basin, equatorial East Africa, Quat. Int, vol.331, pp.20-30, 2014.

A. Shaw, The archaeology of persistent places: the Palaeolithic case of La Cotte de St Brelade, Jersey. Antiquity, vol.90, pp.1437-1453, 2016.
URL : https://hal.archives-ouvertes.fr/halshs-01737618

M. Pope, J. Mcnabb, and C. Gamble, Crossing the human threshold. Dynamic transformation and persistent places during the Middle Pleistocene, 2018.

R. Bernoussi, La faune de vertébrés du Pléistocène moyen de la Grotte des Rhinocéros, Casablanca, Maroc: Approche taphonomique préliminaire, Prehistoire de Casablanca. I -La Grotte des Rhinoce?os (fouilles 1991 et, pp.141-143, 1996.

L. R. Binford, Faunal Remains from Klasies River Mouth, 1984.

Y. M. Lam, X. Chen, and O. M. Pearson, Intertaxonomic Variability in Patterns of Bone Density and the Differential Representation of Bovid, Cervid, and Equid Elements in the Archaeological Record, Am. Antiq, vol.64, pp.343-362, 1999.

R. L. Lyman, Vertebrate taphonomy, 1994.

N. D. Munro, G. Bar-oz, and A. J. Stutz, Aging mountain gazelle (Gazella gazella): refining methods of tooth eruption and wear and bone fusion, J. Archaeol. Sci, vol.36, pp.752-763, 2009.

H. T. Bunn and T. R. Pickering, Methodological recommendations for ungulate mortality analyses in paleoanthropology, Quat. Res, vol.74, pp.388-394, 2010.

P. M. Hitchins, Age determination of the black rhinoceros, Diceros bicornis Linn.) in Zululand. South African. J. Wildl. Res, vol.8, pp.71-80, 1978.

H. Tong, Age profiles of Rhino Fauna from the middle pleistocene nanjing man site, South China -Explained by the Rhino specimens of living species, Int. J. Osteoarchaeol, vol.11, pp.231-237, 2001.

P. Villa and E. Mahieu, Breakage patterns of human long bones, J. Hum. Evol, vol.21, pp.27-48, 1991.

R. J. Blumenschine and M. M. Selvaggio, Percussion marks on bone surface as a new diagnostic of hominid behaviour, Nature, vol.333, pp.763-765, 1988.

, Scientific RepoRtS |, vol.10, p.4761, 2020.

T. R. Pickering and C. P. Egeland, Experimental patterns of hammerstone percussion damage on bones: Implications for inferences of carcass processing by humans, J. Archaeol. Sci, vol.33, pp.459-469, 2006.

D. Vettese, Towards an understanding of hominin marrow extraction strategies: a proposal for a percussion mark terminology, Archaeol. Anthropol. Sci, 2020.
URL : https://hal.archives-ouvertes.fr/hal-02461271

R. J. Blumenschine, C. W. Marean, and S. D. Capaldo, Blind tests of inter-analyst correspondence and accuracy in the identification of cut marks, percussion marks, and carnivore tooth marks on bone surfaces, J. Archaeol. Sci, vol.23, pp.493-507, 1996.

A. K. Behrensmeyer, Taphonomic and Ecologic Information from Bone Weathering, Palaeobiology, vol.4, pp.150-162, 1978.

M. Domínguez-rodrigo, S. De-juana, A. B. Galán, and M. Rodríguez, A new protocol to differentiate trampling marks from butchery cut marks, J. Archaeol. Sci, vol.36, pp.2643-2654, 2009.

A. K. Behrensmeyer, K. D. Gordon, and G. T. Yanagi, Trampling as a cause of bone surface damage and pseudo-cutmarks, Nature, vol.319, pp.768-771, 1986.

C. K. Brain, The Hunters or the Hunted? An Introduction to African Cave Taphonomy, 1981.

L. R. Binford, Bones: ancient men and modern myths, 1981.

G. Haynes, A guide for differentiating mammalian carnivore taxa responsible for gnaw damage to herbivore limb bones, Paleobiology, vol.9, pp.164-172, 1983.

S. L. Olsen and P. Shipman, Surface modification on bone: Trampling versus butchery, J. Archaeol. Sci, vol.15, pp.535-553, 1988.

R. J. Blumenschine, An Experimental Model of the Timing of Hominid and Carnivore Influence on Archaeological Bone Assemblages, J. Archaeol. Sci, vol.15, pp.483-502, 1988.

M. M. Selvaggio and J. Wilder, Identifying the involvement of multiple carnivore taxa with archaeological bone assemblages, J. Archaeol. Sci, vol.28, pp.465-470, 2001.

M. Domínguez-rodrigo and A. Piqueras, The use of tooth pits to identify carnivore taxa in tooth-marked archaeofaunas and their relevance to reconstruct hominid carcass processing behaviours, J. Archaeol. Sci, vol.30, pp.1385-1391, 2003.

T. R. Pickering, M. Domínguez-rodrigo, C. P. Egeland, and C. K. Brain, Beyond leopards: Tooth marks and the contribution of multiple carnivore taxa to the accumulation of the Swartkrans Member 3 fossil assemblage, J. Hum. Evol, vol.46, pp.595-604, 2004.

H. W. Tong, S. Zhang, F. Chen, and Q. Li, Rongements sélectifs des os par les porcs-épics et autres rongeurs: cas de la grotte Tianyuan, un site avec des restes humains fossiles récemment découvert près de Zhoukoudian (Choukoutien), Anthropologie, vol.112, pp.353-369, 2008.

R. Blasco, J. Rosell, J. Fernández-peris, I. Cáceres, and J. M. Vergès, A new element of trampling: an experimental application on the Level XII faunal record of Bolomor Cave, J. Archaeol. Sci, vol.35, pp.1605-1618, 2008.

P. Shipman and J. J. Rose, Cutmark Mimics on Modern and Fossil Bovid Bones, Curr. Anthropol, vol.25, pp.116-117, 1984.

J. De-heinzelin, Environment and behavior of 2.5-million-year-old Bouri hominids. Science (80-.), vol.284, pp.625-629, 1999.

I. Fiore, Taphonomic analysis of the Late Early Pleistocene bone remains from Buia (Dandiero Basin, Danakil depression, Eritrea): Evidence for large mammal and reptile butchering, Riv. Ital. di Paleontol. e Stratigr, vol.110, pp.89-97, 2004.

M. Domínguez-rodrigo, T. R. Pickering, S. Semaw, and M. J. Rogers, Cutmarked bones from Pliocene archaeological sites at Gona, Afar, Ethiopia: Implications for the function of the world's oldest stone tools, J. Hum. Evol, vol.48, pp.109-121, 2005.

R. L. Lyman and . Quantitative, , 2008.

C. K. Brain, Some principles in the interpretation of bone accumulations associated with man. in Human origins, pp.96-116, 1976.

T. D. White, Prehistoric Cannibalism at Mancos 5MTUMR-2346, 1992.

Y. Fernández-jalvo, J. C. Díez, I. Cáceres, and J. Rosell, Human cannibalism in the Early Pleistocene of Europe, J. Hum. Evol, vol.37, pp.591-622, 1999.

Y. Fernández-jalvo and P. Andrews, When humans chew bones, J. Hum. Evol, vol.60, pp.117-123, 2011.

S. M. Bello, P. Saladié, I. Cáceres, A. Rodríguez-hidalgo, and S. A. Parfitt, Upper Palaeolithic ritualistic cannibalism at Gough's Cave, J. Hum. Evol, vol.82, pp.170-189, 2015.

P. Saladié, A. Rodríguez-hidalgo, C. Díez, P. Martín-rodríguez, and E. Carbonell, Range of bone modifications by human chewing, J. Archaeol. Sci, vol.40, pp.380-397, 2013.

R. Grün, H. P. Schwarcz, and J. M. Chadam, ESR dating of tooth enamel: coupled correction for U-uptake and U-series disequilibrium, Nucl. Tracks Radiat. Meas, vol.14, pp.237-241, 1988.

Q. Shao, J. Bahain, C. Falguères, J. M. Dolo, and T. Garcia, A new U-uptake model for combined ESR/U-series dating of tooth enamel, Quat. Geochronol, vol.10, pp.406-411, 2012.

R. Grün, Direct dating of human remains, Yearb. Phys. Anthropol, vol.49, pp.2-48, 2006.

C. Falguères, A 300-600 ka ESR/U-series chronology of Acheulian sites in Western Europe, Quat. Int, pp.293-298, 2010.

J. L. Bischoff, R. J. Rosenbauer, A. Tavoso, and H. De-lumley, A test of uranium-series dating of fossil tooth enamel: Results from Tournal cave, France. Appl. Geochemistry, vol.3, pp.135-141, 1988.

E. Pons-branchu, C. Hillaire-marcel, P. Deschamp, B. Ghaleb, and D. J. Sinclair, Early diagenesis impact on precise U-series dating of deep-sea corals: example of a 100-200-years old Lophelia pertusa sample from the northeast Atlantic, Geochim. Cosmochim. Acta, vol.69, pp.4865-4879, 2005.

Y. Yokoyama and H. V. Nguyen, Direct and non destructive dating of marine sediments, manganese nodules and corals by high resolution gamma-ray spectrometry, Isotope marine chemistry, pp.259-289, 1980.

R. Grün, R. Joannes-boyau, and C. Stringer, Two types of CO2? radicals threaten the fundamentals of ESR dating of tooth enamel, Quat. Geochronol, vol.3, pp.150-172, 2008.

R. Grün, The DATA program for the calculation of ESR age estimates on tooth enamel, Quat. Geochronol, vol.4, pp.231-232, 2009.

R. Grün and O. Katzenberger-apel, An alpha irradiator for ESR dating, Anc. TL, vol.12, pp.35-38, 1994.

B. J. Brennan, W. J. Rink, E. L. Mcguirl, H. P. Schwarcz, and W. V. Prestwich, One Group"' theory and the Rosy ESR dating software, Radiat. Meas, vol.27, pp.307-314, 1997.

G. Guérin, N. Mercier, R. Nathan, G. Adamiec, and Y. Lefrais, On the use of the infinite matrix assumption and associated concepts: A critical review, Radiat. Meas, vol.47, pp.778-785, 2012.

J. Bahain, Y. Yokoyama, C. Falguères, and M. N. Sarcia, ESR dating of tooth enamel: a comparison with K-Ar dating, Quat. Sci. Rev, vol.11, pp.245-250, 1992.

J. J. Hublin, New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens, Nature, vol.546, pp.289-292, 2017.

, Scientific RepoRtS |, vol.10, p.4761, 2020.

V. D. , R. G. , A. M. , J. P. , and J. P. , It has received funding from the Aquitaine Region through the Origines projects, the Collège de France, the Muséum National d'Histoire Naturelle in Paris, and the University of Bordeaux (France), Lithics were studied by