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A. , B. Brou, and D. Coulibaly, This work was supported by RIPIECSA-Côte d'Ivoire Project (Recherche Interdisciplianaire et Participative sur les Interactions entre les Ecosystèmes, le Climat et les Sociétés en Afrique de l'Ouest). A.B.N'.D. thanks the BDI fellowship programme of CNRS (Centre National de la Recherche Scientifique) for scholarship. We thank François N'Guessan, Honoré Kounan, ACKNOWLEDGEMENTS. We thank Sébastien Barot for comments on the manuscript

, , 2016.