G. Bell, Ecology-neutral macroecology, Science, vol.293, pp.2413-2418, 2001.

S. P. Hubbell, Neutral theory in community ecology and the hypothesis of functional equivalence, Funct. Ecol, vol.19, pp.166-172, 2005.

P. B. Adler, J. Hillerislambers, and J. M. Levine, A niche for neutrality, Ecol. Lett, vol.10, pp.95-104, 2007.

J. J. Wiens, R. A. Pyron, and D. S. Moen, Phylogenetic origins of local-scale diversity patterns and the causes of Amazonian megadiversity, Ecol. Lett, vol.14, pp.643-652, 2011.

M. Wolf, G. S. Van-doorn, O. Leimar, and F. J. Weissing, Life-history trade-offs favour the evolution of animal personalities, Nature, vol.447, pp.581-584, 2007.

A. T. Vanak, D. Fortin, M. Thaker, M. Ogden, C. Owen et al., Moving to stay in place: behavioral mechanisms for coexistence of African large carnivores, Ecology, vol.94, pp.2619-2631, 2013.

O. Spiegel, S. T. Leu, C. M. Bull, and A. Sih, What's your move? Movement as a link between personality and spatial dynamics in animal populations, Ecol. Lett, vol.20, pp.3-18, 2017.

G. Péron, The time frame of home-range studies: from function to utilization, Biol. Rev. In press, 2019.

J. H. Grobler, Host selection and species preference of the red-billed oxpecker Buphagus erythrorhynchus in the Kruger National Park, vol.23, pp.89-97, 1980.

I. M. Stutterheim and K. Panagis, Roosting behaviour and host selection of oxpeckers (Aves: Buphaginae) in Moremi Wildlife Reserve, 1985.

S. Afr, J. Zool, vol.20, pp.237-240

K. Hustler, Host preference of oxpeckers in the Hwange National Park, Zimbabwe. Afr. J. Ecol, vol.25, pp.241-245, 1987.

W. D. Koenig, Host preferences and behaviour of oxpeckers: co-existence of similar species in a fragmented landscape, Evol. Ecol, vol.11, pp.91-104, 1997.

C. L. Nunn, V. O. Ezenwa, C. Arnold, and W. D. Koenig, Mutualism or parasitism? Using a phylogenetic approach to characterize the oxpecker-ungulate relationship, Evolution, vol.65, pp.1297-1304, 2011.

M. Ndlovu and L. Combrink, Feeding preferences of oxpeckers in Kruger National Park, South Africa. Koedoe, vol.57, pp.1-6, 2015.

P. Mikula, J. Hadrava, T. Albrecht, and P. Tryjanowski, Large-scale assessment of commensalisticmutualistic associations between African birds and herbivorous mammals using internet photos, PeerJ, vol.6, p.4520, 2018.

C. H. Fry, S. Keith, and E. K. Urban, The birds of Africa, vol.6, 2000.

A. J. Hall-martin, Range expansion of the yellowbilled oxpecker Buphagus africanus into the, 1987.

P. Kruger-national, South Africa. Koedoe, vol.30, pp.121-132

M. Begon, J. L. Harper, and C. R. Tounsend, Ecology: individuals, populations and communities, 1986.

R. Arlettaz, N. Perrin, and J. Hausser, Trophic resource partitioning and competition between the two sibling bat species Myotis myotis and Myotis blythii, J. Anim. Ecol, vol.66, pp.897-912, 1997.

R. Gagnon, C. T. Mabika, and C. Bonenfant, Distribution and density of oxpeckers on giraffes in, 2019.

P. Chesson, Mechanisms of maintenance of species diversity, Annu. Rev. Ecol. Syst, vol.31, pp.343-366, 2000.

P. Marjoram, J. Molitor, V. Plagnol, and S. Tavare, Markov chain Monte Carlo without likelihoods, Proc. Natl Acad. Sci. USA, vol.100, pp.324-339, 2003.

M. J. White, Segregation and diversity measures in population distribution, Popul. Index, vol.52, pp.198-221, 1986.

R. A. Noske, Co-existence of three species of treecreepers in north-eastern, New South Wales. Emu, vol.79, pp.120-128, 1979.

J. Martin and J. Thibault, Coexistence in Mediterranean warblers: ecological differences or interspecific territoriality?, J. Biogeogr, vol.23, p.20190548, 1996.

R. B. Root, The niche exploitation pattern of the blue-gray gnatcatcher, Ecol. Monogr, vol.37, pp.317-350, 1967.

C. M. Herrera, Ecological aspects of heterospecific flocks formation in a Mediterranean passerine bird community, Oikos, vol.33, pp.85-96, 1979.

T. Hino, Mutualistic and commensal organization of avian mixed-species foraging flocks in a forest of western Madagascar, J. Avian Biol, vol.29, pp.17-24, 1998.

M. S. Palmer and C. Packer, Giraffe bed and breakfast: camera traps reveal Tanzanian yellow-billed oxpeckers roosting on their large mammalian hosts, Afr. J. Ecol, vol.56, pp.882-884, 2018.

G. Péron, Multicontinental community phylogenetics of avian mixed-species flocks reveal the role of the stability of associations and of kleptoparasitism, Ecography, vol.40, pp.1267-1273, 2017.

D. Grünbaum and R. R. Veit, Black-browed albatrosses foraging on Antarctic krill: densitydependence through local enhancement?, Ecology, vol.84, pp.3265-3275, 2003.

C. Mettke-hofmann, M. Wink, M. Braun, and H. Winkler, Residency and a broad feeding spectrum are related to extensive spatial exploration in parrots, Behav. Ecol, vol.23, pp.1365-1371, 2012.

M. D. Sorenson, The functional significance of parasitic egg laying and typical nesting in redhead ducks: an analysis of individual behavior, Anim. Behav, vol.42, pp.80122-80130, 1991.

G. Péron and D. N. Koons, Integrated modeling of communities: parasitism, competition, and demographic synchrony in sympatric ducks, Ecology, vol.93, pp.2456-2464, 2012.

G. Péron and P. Crochet, Edge effect and structure of mixed-species bird flocks in an Afrotropical lowland forest, J. Ornithol, vol.150, pp.585-599, 2009.

C. J. Stutterheim and R. K. Brooke, Past and present ecological distribution of the yellowbilled oxpecker in South Africa, S. Afr. J. Zool, vol.16, pp.44-49, 1981.

A. Robertson and A. M. Jarvis, Oxpeckers in northeastern Namibia: recent population trends and the possible negative impacts of drought and fire, Biol. Conserv, vol.92, issue.99, pp.69-74, 2000.

N. Diplock, Large mammal declines and the incipient loss of mammal-bird mutualisms in an African savanna ecosystem, PLoS ONE, vol.13, 2018.

G. Péron, C. Bonenfant, R. Gagnon, and C. T. Mabika, Data from: The two oxpecker species reveal the role of movement rates and foraging intensity in species coexistence, Dryad Digital Repository, 2019.

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